Founder: Northern (Arctic) Federal University named after M.V. Lomonosov

Editorial office address: Russian Federation, 163002, Arkhangelsk, Naberezhnaya Severnoy Dviny 17, office 1410a

Phone: (818-2) 21-61-00(15-33)



Manuscript preparation

Версия для печати


Title: The title should be in a sentence case (only scientific, geographic or person names should be with a first capital letter, i.e. Elater ferrugineus L., Germany, etc.), and should include an accurate, clear and concise description of the reported work, avoiding abbreviations. The higher taxa within the title should be separated with commas and not with a semicolon, e.g.: (Coleoptera, Elateridae, Elaterini).

Authors and Affiliations: Provide the complete names of all authors, and their addresses for correspondence, including e.g., institutional affiliation (e.g. university, institute), location (street, boulevard), city, state/province (if applicable), and country. One of the authors should be designated as the corresponding author. It is the corresponding author's responsibility to ensure that the author list, and the individual contributions to the study are accurate and complete. If the article has been submitted on behalf of a consortium, all consortium members and their affiliations should be listed after the Acknowledgements section.

Abstract and Keywords: Please have your abstract and keywords ready for input into the submission module. Keywords should be in alphabetical order and ideally differ from the words used in the title.

Body Text: All papers should be in grammatically correct English. Non-native English speaking authors are required to have their manuscripts checked by a native English speaker prior to submission. Use either British/Commonwealth or American English provided that the language is consistent within the paper. A manuscript must be written with precision, clarity, and economy. The voice - active or passive - and the tense used should be consistent throughout the manuscript. Avoid the use of parenthetical comments and italics or bold for emphasis. This journal discourages the use of quotation marks except for direct quotations, words defined by the author, and words used in unusual contexts. Short quotations should be embedded in the text and enclosed in double quotation marks ("). Long quotations should be on a separate line, italicized, but without quotation marks. Single quotation marks are to be used only for a quotation that occurs within another quotation.

Spacing, Fonts, and Page Numbering: Single-space all material (text, quotations, figure legends, tables, references, etc.). Separate paragraphs with a blank line. Use a 12-point font (preferably Times New Roman or Arial).

Capitals: First capital letters should be used only in the beginning of a sentence, in proper names and in headings and subheadings, as well as to indicate tables, graphs and figure/s within the text. Software programmes should be written with capital letters (e.g., ANOVA, MANOVA, PAUP).

Italicization/Underlining: Scientific names of species and genera, long direct quotations and symbols for variables and constants (except for Greek letters), such as p, F, U, T, N, r, but not for SD (standard deviation), SE (standard error), DF (degrees of freedom) and NS (non significant) should be italicized. These symbols in illustrations and equations should be in italics to match the text. Italics should not be used for emphasis, and not in abbreviations such as e.g., i.e., et al., etc., cf. Underlining of any text is not acceptable.

Abbreviations: Abbreviations should be followed by ‘.' (full stop or period; for instance: i.e., e.g., cf., etc.). Note that you shouldn't add a full stop at the end of abbreviated words if the last letter of the abbreviation is the same as the last letter of the full word. For example, you should abbreviate "Eds", "Dr", "Mr" without full stop at the end. All measures, for instance mm, cm, m, s, L, should be written without full stop.

On the use of dashes: (1) Hyphens are used to link words such as personal names, some prefixes and compound adjectives (the last of which vary depending on the style manual in use) (2) En-dash or en-rule (the length of an 'n') is used to link spans. In the context of our journal en-dash should be used to link numerals, sizes, dates and page numbers (e.g., 1977–1981; figs 5–7; pp. 237–258); geographic or name associations (Murray–Darling River; a Federal–State agreement); and character states combinations such as long–pubescent or red–purple. (3) Em-dash or em-rule (the length of an 'm') should be used rarely, only for introducing a subordinate clause in the text that is often used much as we use parentheses. In contrast to parentheses an em-dash can be used alone. En-dashes and em-dashes should not be spaced.

Footnotes: Avoid footnotes in the body text of the manuscript. It is always possible to incorporate the footnote into the main text by rewording the sentences, which greatly facilitates reading. Additionally, footnotes are not always handled well by the journal software, and their usage may cause a failure of submission. Footnotes are acceptable only below tables; instead of numbers, please use (in order): †, ‡, §, |, ¶, #, ††, ‡‡, §§, ||, ¶, ##.

Geographical coordinates: It is strongly recommended to list geographical coordinates as taken from GPS or online gazetteer, or georeferencer. Geographical coordinates must be listed in one of the following formats:

Definition: The locality consists of a point represented by coordinate information in the form of latitude and longitude. Information may be in the form of

  • Degrees, Minutes and Seconds (DMS),
  • Degrees and Decimal Minutes (DDM), or
  • Decimal Degrees (DD).

Records should also contain a hemisphere (E or W and N or S) or, with Decimal Degrees, minus (–) signs to indicate western and/or southern hemispheres.


  • Example 1: 36° 31' 21" N; 114° 09' 50" W (DMS)
  • Example 2: 36° 31.46’N; 114° 09.84’W (DDM)
  • Example 3: 36.5243° S; 114.1641° W (DD)
  • Example 4: −36.5243; −114.1641 (DD using minus signs to indicate southern and western hemispheres)

Note on accuracy: Because GPS units are very commonly used today to record latitude/longitude, many authors simply give the GPS readings for their localities. However, these readings are much too accurate. For example, a GPS unit might give the latitude in decimal seconds as 28°16'55.87"N. Since one second of latitude is about 30 m on the ground, the second figure after the decimal in 55.87 represents 30 cm, yet a typical handheld GPS unit is only accurate at best to a few metres.

We therefore recommend two ways to report GPS-based locations. If you give the GPS reading without rounding off, make sure you include an uncertainty figure as a context for the over-accurate GPS reading. We recommend the Darwin Core definition of uncertainty (

"The horizontal distance (in meters) from the given decimalLatitude and decimalLongitude describing the smallest circle containing the whole of the Location."

If you only give the GPS reading, please round it off to an implied precision appropriate to the error in the measurement, or to the extent of the area sampled. We suggest rounding off

  • to the nearest second in degree-minute-second format (28°16'56"N), which implies roughly ± 25-30 m at middle latitudes
  • to four decimal places in decimal degree format (28.2822°N), which implies roughly ± 10-15 m at middle latitudes
  • to two decimal places in decimal minute format (28°16.93'N), which implies roughly 15-20 m at middle latitudes

Altitude: Many GPS users simply record the elevation given by their GPS unit. However, GPS elevation is NOT the same as elevation above sea level. GPS units record the elevation above a mathematical model of the earth's surface. The difference between this elevation and elevation above sea level can be tens of metres. In any case, the accuracy of a GPS elevation is often the same as the usual accuracy in horizontal position, so a GPS elevation such as '753 m' is much too accurate and should be rounded off to 'ca 750 m'.

We strongly recommend the use of Example 2 (the DDM format). The other three are also possible but will be recalculated to DDM during the process of online mapping from the HTML version of the paper.

The only restriction on format is in creating a KML (Keyhole Markup Language) file. KML latitudes and longitudes must be in the DD format shown above in Example 4.

Please also consider submitting a table of localities with your manuscript, either as a spreadsheet or in CSV text format. By doing so you will make your specimen localities much more easily available for use in biodiversity databases and geospatial investigations. The geospatial table will be put online as supplementary material for your paper. A minimum table will have three fields: species (or subspecies) name, latitude and longitude. A full table will have the same data for each specimen lot as appears in the text of your paper. Please check latitude/longitude carefully for each entry.

Units: Use the International System of Units (SI) for measurements. Consult Standard Practice for Use of the International System of Units (ASTM Standard E−380−93) for guidance on unit conversions, style, and usage.

Statistics: Use leading zeroes with all numbers, including probability values (e.g., P < 0.001). For every significant F−statistic reported, provide two df values (numerator and denominator). Whenever possible, indicate the year and version of the statistical software used.

Web (HTML) links: Authors are encouraged to include links to other Internet resources in their article. This is especially encouraged in the reference section. When inserting a reference to a web-page, please include the http:// portion of the web address.

Supplementary files: Larger datasets can be uploaded separately as Supplementary Files. Tabular data provided as supplementary files can be uploaded as an Excel spreadsheet (.xls), as an OpenOffice spreadsheets (.ods) or comma separated values file (.csv). As with all uploaded files, please use the standard file extensions.

Headings and subheadings: Main headings: The body text should be subdivided into different sections with appropriate headings. Where possible, the following standard headings should be used: Introduction, Methods, Results, Discussion, Conclusions, Acknowledgements, References. These headings need to be in bold font on a separate line and start with a first capital letter. Please do not number headings or subheadings.

  • Introduction − The motivation or purpose of your research should appear in the Introduction, where you state the questions you sought to answer, and then provide some of the historical basis for those questions.
  • Methods − Provide sufficient information to allow someone to repeat your work. A clear description of your experimental design, sampling procedures, and statistical procedures is especially important in papers describing field studies, simulations, or experiments. If you list a product (e.g., animal food, analytical device), supply the name and location of the manufacturer. Give the model number for equipment used. Supply complete citations, including author (or editor), title, year, publisher, and version number, for computer software mentioned in your article.
  • Results − Results should be stated concisely and without interpretation.
  • Discussion − Focus on the rigorously supported aspects of your study. Carefully differentiate the results of your study from data obtained from other sources. Interpret your results, relate them to the results of previous research, and discuss the implications of your results or interpretations. Point out results that do not support speculations or the findings of previous research, or that are counter-intuitive. You may choose to include a Speculation subsection in which you pursue new ideas suggested by your research, compare and contrast your research with findings from other systems or other disciplines, pose new questions that are suggested by the results of your study, and suggest ways of answering these new questions.
  • Conclusion −This should state clearly the main conclusions of the research and give a clear explanation of their importance and relevance. Summary illustrations may be included.
  • References − The list of References should be included after the final section of the main article body. A blank line should be inserted between single-spaced entries in the list. Authors are requested to include links to online sources of articles, whenever possible! 

Where possible, the standard headings should be used in the order given above. Additional headings and modifications are permissible.

Subordinate headings: Subordinate headings (e.g. Field study and Simulation model or Counts, Measurements and Molecular analysis), should be left-justified, italicized, and in a regular sentence case. All subordinate headings should be on a separate line.


Figures and illustrations are accepted in the following image file formats:

  • EPS (preferred format for diagrams)
  • TIFF (at least 300dpi resolution, with LZW compression)
  • PNG (preferred format for photos or images)
  • JPEG (preferred format for photos or images)
  • GIF
  • BMP
  • SVG

Figure legends: All figures should be referenced consecutively in the manuscript; legends should be listed consecutively immediately after the References. For each figure, the following information should be provided: Figure number (in sequence, using Arabic numerals − i.e. Figure 1, 2, 3 etc.); short title of figure (maximum 15 words); detailed legend, up to 300 words.

Illustrations of measurable morphological traits should bear mute scale bars, whose real size is to be given in the figure captions.

Please note that it is the responsibility of the author(s) to obtain permission from the copyright holder to reproduce figures or tables that have previously been published elsewhere.

On the use of Google Maps
Please do NOT use maps produced by Google Earth and Google Maps in your publications, as these are subject of copyright! Here is an excerpt from Google Maps/Earth Additional Terms of Service:
Restrictions on Use. Unless you have received prior written authorization from Google (or, as applicable, from the provider of particular Content), you must not: (a) copy, translate, modify, or make derivative works of the Content or any part thereof; (b) redistribute, sublicense, rent, publish, sell, assign, lease, market, transfer, or otherwise make the Products or Content available to third parties; (c) reverse engineer, decompile or otherwise attempt to extract the source code of the Service or any part thereof, unless this is expressly permitted or required by applicable law; (d) use the Products in a manner that gives you or any other person access to mass downloads or bulk feeds of any Content, including but not limited to numerical latitude or longitude coordinates, imagery, and visible map data; (e) delete, obscure, or in any manner alter any warning or link that appears in the Products or the Content; or (f) use the Service or Content with any products, systems, or applications for or in connection with (i) real time navigation or route guidance, including but not limited to turn-by-turn route guidance that is synchronized to the position of a user's sensor-enabled device; or (ii) any systems or functions for automatic or autonomous control of vehicle behavior; (g) use the Products to create a database of places or other local listings information.

Tables: Each table should be numbered in sequence using Arabic numerals (i.e. Table 1, 2, 3 etc.). Tables should also have a title that summarizes the whole table, maximum 15 words. Detailed legends may then follow, but should be concise.

Small tables can be embedded within the text, in portrait format (note that tables on a landscape page must be reformatted onto a portrait page or submitted as additional files). These will be typeset and displayed in the final published form of the article. Such tables should be formatted using the 'Table object' in a word processing program to ensure that columns of data are kept aligned when the file is sent electronically for review. Do not use tabs to format tables or separate text. All columns and rows should be visible, please make sure that borders of each cell display as black lines. Colour and shading should not be used; neither should commas be used to indicate decimal values. Please use a full stop to denote decimal values (i.e., 0.007 cm, 0.7 mm).

Larger datasets can be uploaded separately as Supplementary Files. Tabular data provided as supplementary files can be uploaded as an Excel spreadsheet (.xls), as an OpenOffice spreadsheets (.ods) or comma separated values file (.csv). As with all uploaded files, please use the standard file extensions.


Online publishing allows an author to provide datasets, tables, video files, or other information as supplementary information, greatly increasing the impact of the submission. Uploading of such files is possible in Step 6 of the submission process.

The maximum file size for each Supplementary File is 20 MB.

The Supplementary Files will not be displayed in the printed version of the article but will exist as linkable supplementary downloadable files in the online version.

While submitting a supplementary file the following information should be completed:

  • File format (including name and a URL of an appropriate viewer if format is unusual)
  • Title of data
  • Description of data

All supplementary files should be referenced explicitly by file name within the body of the article, e.g. 'See supplementary file 1: Movie 1" for the original data used to perform this analysis.

Ideally, the supplementary files should not be platform-specific, and should be viewable using free or widely available tools. Suitable file formats are:

For supplementary documentation:

  • PDF (Adobe Acrobat)

For animations:

  • SWF (Shockwave Flash)

For movies:

  • MOV (QuickTime)
  • MPG (MPEG)

For datasets:

  • XLS (Excel spreadsheet)
  • CSV (Comma separated values)
  • ODS (OpenOffice spreadsheets)

As for images, file names should be given in the standard file extensions. This is especially important for Macintosh users, since the Mac OS does not enforce the use of standard file extensions. Please also make sure that each additional file is a single table, figure or movie (please do not upload linked worksheets or PDF files larger than one sheet).


Citations within the text: Before submitting the manuscript, please check each citation in the text against the References and vice-versa to ensure that they match exactly. Citations in the text should be formatted as follows: Smith (1990) or (Smith 1990), Smith et al. (1998) or (Smith et al. 1998) and (Smith et al. 1998, 2000, Brock and Gunderson 2001, Felt 2006).

References: It is important to format the references properly, because all references will be linked electronically as completely as possible to the papers cited. It is desirable to add a DOI (digital object identifier) number for either the full-text or title and abstract of the article as an addition to traditional volume and page numbers. If a DOI is lacking, it is recommended to add a link to any online source of an article. Please use the following style for the reference list (or download the Pensoft EndNote style): here

Published Papers:
Polaszek A, Alonso-Zarazaga M, Bouchet P, Brothers DJ, Evenhuis NL, Krell FT, Lyal CHC, Minelli A, Pyle RL, Robinson N, Thompson FC, van Tol J (2005) ZooBank: the open-access register for zoological taxonomy: Technical Discussion Paper. Bulletin of Zoological Nomenclature 62: 210-220.

Accepted Papers:
Same as above, but ''in press'' appears instead the year in parentheses.

Electronic Journal Articles:
Mallet J, Willmott K (2002) Taxonomy: renaissance or Tower of Babel? Trends in Ecology and Evolution 18 (2): 57-59. doi: 10.1016/S0169-5347(02)00061-7.

Paper within conference proceedings:
Orr AG (2006) Odonata in Bornean tropical rain forest formations: Diversity, endemicity and applications for conservation management. In: Cordero Rivera A (Ed) Forest and Dragonflies. Fourth WDA International Symposium of Odonatology, Pontevedra (Spain), July 2005. Pensoft Publishers, Sofia-Moscow, 51-78.

Book chapters:
Mayr E (2000) The biological species concept. In: Wheeler QD, Meier R (Eds) Species Concepts and Phylogenetic Theory: A Debate. Columbia University Press, New York, 17-29.

Goix N, Klimaszewski J (2007) Catalogue of Aleocharine Rove Beetles of Canada and Alaska. Pensoft Publishers, Sofia-Moscow, 166 pp.

Book with institutional author:
International Commission on Zoological Nomenclature (1999) International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature.

PhD thesis:
Dalebout ML (2002) Species identity, genetic diversity and molecular systematic relationships among the Ziphiidae (beaked whales). PhD thesis, Auckland, New Zealand: University of Auckland.

BBC News: Island leopard deemed new species

Citations of Public Resource Databases: It is highly recommended all appropriate datasets, images, and information to be deposited in public resources. Please provide the relevant accession numbers (and version numbers, if appropriate). Accession numbers should be provided in parentheses after the entity on first use. Examples of such databases include, but are not limited to:

Providing accession numbers to data records stored in global data aggregators allows us to link your article to established databases, thus integrating it with a broader collection of scientific information. Please hyperlink all accession numbers through the text or list them directly after the References in the online submission manuscript.

All journal titles should be spelled out completely and should NOT be italicized.

Provide the publisher's name and location when you cite symposia or conference proceedings; distinguish between the conference date and the publication date if both are given. Do not list abstracts or unpublished material in the References. They should be quoted in the text as personal observations, personal communications, or unpublished data, specifying the exact source, with date if possible. When possible, include URLs for articles available online through library subscription or individual journal subscription, or through large international archives, indexes and aggregators, e.g., PubMedCentral, Scopus, CAB Abstracts, etc. URLs for pdf articles that are posted on personal websites only should be avoided.

Authors are encouraged to cite in the References list the publications of the original descriptions of the taxa treated in their manuscript.





Journal of Medical and Biological

Vestnik NArFU.
Series "Humanitarian
and Social Sciences"

Forest Journal 
Лесной журнал 

Arctic and North